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Population Dynamics with Two Competing Species

stream plot
stability
growth rate
α
1
1
α
2
0.5
self-inhibition
β
1
1
β
2
0.5
interaction
γ
1
0.6
γ
2
0.75
Consider two types of fish in a pond that do not prey on one another but compete for the available food. The governing equations are:
x
t
=x(
α
1
-
β
1
x-
γ
1
y)=x
Δ
1
(x,y)
,
y
t
=y(
α
2
-
β
2
y-
γ
2
x)=y
Δ
2
(x,y)
.
Restrict the discussion to the first quadrant, since
x
and
y
are species populations.
There are up to four steady states in the first quadrant:
S
1
=(0,0)
,
S
2
=0,
α
2
β
2
(or
Δ
2
(x,y)=0
),
S
3
=
α
1
β
2
,0
(or
Δ
1
(x,y)=0
),
S
4
=
α
1
β
2
-
α
2
γ
1
β
1
β
2
-
γ
1
γ
2
,
α
2
β
1
-
α
1
γ
2
β
1
β
2
-
γ
1
γ
2
(or
Δ
1
(x,y)=0
and
Δ
2
(x,y)=0
).
S
1
,
S
2
, and
S
3
are always in the first quadrant.
S
4
pertains when both species are present in the first quadrant if
α
1
β
2
-
α
2
γ
1
β
1
β
2
-
γ
1
γ
2
>0
and
α
2
β
1
-
α
1
γ
2
β
1
β
2
-
γ
1
γ
2
>0
.
Vary the growth rate coefficients
α
1
,
α
2
, the self-inhibition parameters
β
1
,
β
2
, and the interaction parameters
γ
1
,
γ
2
. This Demonstration gives the corresponding stream plot.
The steady states and nullclines
Δ
1
(x,y)=0
and
Δ
2
(x,y)=0
(blue and brown) are shown in the plot.
From the snapshots,
S
1
is an unstable node,
S
2
and
S
3
are either saddle points or stable nodes, and
S
4
is either a saddle point or a stable node.
The linearized analysis (see the stability tab) confirm these conclusions. Indeed, for
S
1
, both eigenvalues are positive. For
S
2
,
S
3
, and
S
4
, there are either two negative eigenvalues or two real eigenvalues of opposite sign.
Coexistence (i.e., steady-state
S
4
) is possible only if the self-inhibition term dominates the interaction term (
β
1
,
β
2
>
γ
1
,
γ
2
).
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